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Upon learning of Rui's death, Muzan kills the remaining Lower Ranks due to their constant failures; he spares Lower One Enmu and sends him on a mission to kill Tanjiro. After a battle against Enmu in a passenger train, Tanjiro kills him with Kyojuro's support. However, Upper Three Akaza appears and battles Kyojuro, who dies from his wounds as the demon escapes. Months later, the group is sent to Yoshiwara with Tengen, where they face off and kill the Upper Six siblings Gyutaro and Daki. Tengen retires from the Corps after suffering extensive injuries. During another battle at the Swordsmith Village, where the Corps' weapons are forged, Tanjiro meets Genya Shinazugawa, Sanemi's younger brother, who has the mysterious ability to temporarily gain demonic powers after eating their flesh. With the help of Muichiro and Mitsuri, the group manage to kill Upper Five Gyokko and Upper Four Hantengu. In the aftermath, they realize that Nezuko is invulnerable to sunlight. She becomes Muzan's prime target as he has long sought a way to overcome the sun and transform into the ultimate being. Tanjiro also learns his Hinokami Kagura descends from "Sun Breathing", the first Breathing Style created by Yoriichi Tsugikuni, the most powerful Demon Slayer in history, and develops it to fight Muzan. Kagaya forecasts Muzan's intentions and enacts a strict training regimen to prepare the Corps for the upcoming battle.